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I. Community structure refers to the physical appearance, species diversity and abundance, and ecological relationships in an ecosystem (p.
144)
A. biodiversity varies across different ecosystems
1. types of biodiversity: ecological, species, genetic, functional (Miller p. 62)
2. in general, diversity increases as you move toward the equator
3. diversity also varies with depth in water
4. pollution has a significant effect on diversity in aquatic systems (such as streams and the Chesapeake)
B. island biodiversity varies with island size and distance to the mainland
1. number of species on an island depends on rates of immigration and extinction
2. larger islands have more species
3. nearer island have more species
4. research supports these theoretical projections
- 90% - 50% rule
C. types of species ecological roles
1. species can be described by their mode of obtaining energy (predator, decomposer, etc) – see chap 3
2. non-native species cause problems (more on this later)
3. indicator species are particularly sensitive to environmental problems (amphibians are a good example; also certain stream insects)
4. umbrella species: A recent article in National Parks calls the Greater Prairie Chicken an umbrella species , because each type of habitat required by this bird also serves as important habitat for many other species, including deer and other prairie birds
5. are all species vitally important? Compare the rivet and redundancy models
6. keystone species have a disproportionate effect on the rest of the ecosystem
a. top predators such as wolves, cougars, or grizzly bears
- Do cougars still exist in the eastern US? check out these sites: Michigan cougar | save the cougar | Eastern Puma Research | Eastern Cougar
- study suggests that predators exert "top-down control" on ecosystem structure
- learn more about the "top-down control" from Where the Wild Things Were
- the effects of wolves in Yellowstone | and a video
- Ecological effects of cougars
- This lab in Wisconsin documents the effects of over-browsing by deer
b. ecosystem engineers such as elephants, beavers, buffalo, prairie dogs, and even ants
- more on prairie dogs
c. providers of important services, such as pollinators | more here - match pollinators with their flowers | Another pollinators page
- examples from the Sonoran desert
- the starfish Pisaster is a classic example from rocky tidal zones of the western North America coast
- salmon are a keystone species in the northwest - even for the forests!!
- cottonwood trees
- hammerhead/tiger sharks
- more on wolves
- flying squirrels
- American Chestnuts
d. Miller suggests that the term "Foundation Species" is broader - see p. 149
D. competition
1. intraspecific competition (within the same species) leads to natural selection, but does not affect community structure
2. interspecific competition refers to competition between different species for resources such as food and habitat
3. interference competition individuals chase each other away or otherwise physically exclude them
4. exploitation competition individual vary in how efficiently they use a resource
5. the competitive exclusion principle states that two species cannot occupy the same niche (classic experiment with mussels - described in class)
6. species reduce competition through resource partitioning - fig. 7-6 and 7-7 and also 4-7: they use different parts of the same habitat or forage at different times
E. predator-prey relationships – fig. 7-8
- deer have become a major pest in suburban areas because of lost predators. Check out local websites on controlling deer: Fairfax County | Montgomery County
- check out the slide show on wolves
- simulation of predator-prey relationships using wolves and rabbits
- camouflage and warning coloration are adaptations to predator/prey relationships
F. non-predatory ecological relationships
1. interactions between species can be positive, negative, or neutral
- overview of the types
- these relationships are examples of co-evolution | more here on co-evolution
2. parasitism is a relationship in which one species benefits and the other is harmed
- parasites and the red queen hypothesis -- running just to stand still
- derive nutrients and energy from the blood/body fluids of another organism (note examples)
- Can have very complicated life cycles often two or more hosts (example deer ticks)
- the world of parasites -- click on a map to find images and information of parasites in a location
- parasite images/drawings (Ohio State)
- Cryptosporidium and Giardia are parasites found in water -- even in the wilderness!!
- graphic images of parasites and their life cycles
- the national parasite collection has some images -- mostly microscopic
- pictures of leeches
- some images from National Geographic
3. mutualism is a relationship in which both species benefit (fig. 7-9)
a. symbiosis organisms living on or in each other
i. lichens
- lichenland has some more pictures
- a professor in Wisconsen has created a list of fungi of the month
ii. myccorhizal fungi
- pictures from the microbe zoo
- additional pictures at the University of Guelph
- still more pictures
iii. termites
b. non-symbiotic
i. herbivores and grasses
ii. dispersal of fruits by birds and small mammals
- flowers display many interesting designs adapted to pollination
- the forgotten pollinators -- many are going extinct or getting rare!
iv. cleaner fish
v. animals that are normally enemies will sometimes form mutual relationships
4. commensalism - one organism benefits and the other is not harmed. (fig 7-10)
a. ephiphytes (distinguish from parasitic plants)
b. barnacles
c. remora and shark
II. Ecological succession
A. Primary succession
1. Occurs in an area previously unoccupied by organisms
2. Examples volcanic islands, places exposed by retreating glaciers, etc.
- Glacier Bay National Park has glaciers that have retreated dramatically in the past 200 years (more on Glacier Bay) | check out satellite images and animations of the changes that have taken place | more satellite images and lots of other info
- vegetation at Mt. St. Helens is growing back after the 1980 eruption | wildlife have also returned
- occasionally a flood can be severe enough to expose rock. One example is found at the spillway of a dam near DeMoines, Iowa | More here | And here
3. example of typical progression shown for Isle Royale in fig. 7-11
B. Secondary Succession
1. Occurs in areas that have been disturbed by some natural phenomenon
2. examples of disturbances: fire, storms, floods, agriculture
C. Examples:
1. field to forest (fig. 7-12)
2. pond to bog to meadow
3. fire-disturbed areas
4. sand dunes
D. Climax communities
1. type of climax community is determined by local climate, soil, rainfall, etc.
2. Succession is not necessarily a predictable event, and some ecologists prefer the term “mature” to “climax” – see discussion p. 157-158
Ecosystems adapted to fire